Edmund Rice Homestead at East Sudbury
Edmund Rice (1638) Association

Rice Family Y-DNA Project

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Highlighted technical terms are explained in the Glossary. Note: a term may be used many times but is highlighted only the first time within each section.

Last updated: 2008 Mar 30


 
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Project Summary


 

Table of Contents


 
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Introduction

The Edmund Rice (1638) Association has a project underway seeking to discover the ancestors of Edmund Rice and other Rice and Royce families. (There was a tendency in past centuries to regard these two spellings as interchangeable.) Y-chromosome DNA analysis offers exciting opportunities to learn more about early family roots. With the assistance of three genetics testing labs, we compared the DNA of many male-line descendants of Edmund Rice of Sudbury and Marlborough and reconstructed the genetic "fingerprint" or haplotype of our immigrant ancestor (see Table 1). For the details of how we did this, see the article entitled "How We Obtained the Rice Haplotype" in our newsletter.

Knowing the haplotypes of Edmund Rice and several other progenitors, we can now invite all Rice/Royce males to compare your DNA against Edmund's and against each other's. For those of you who wonder whether you may be his descendants, such a comparison can help to investigate that possibility. A match with Edmund's haplotype will confirm that you are indeed related (though not necessarily a descendant) and will encourage and aid you in further genealogical research to discover your Rice ancestral line. We have on line a portion of the Rice family history (Edmund and five generations of his descendants) to help in finding the connection if you do match. On the other hand, a big difference from Edmund would indicate you are not related to him at all, but might reveal a similarity to other Rices who are related instead. For male Rices who already have reason to believe you are not Edmund Rice's descendants, the comparison may reveal whether or not the separate Rice families have some connection back in the British Isles. In either case, it should be clear that some conventional genealogical research will be needed in order to get the most out of the DNA results. Table 1, Table 2, and Table 3 below have all the DNA results we have obtained to date.

You, too, can participate. We have arranged with FamilyTree DNA (FTDNA) to offer a 12-locus DNA analysis for a reduced rate of $99 to those who join our project. (Note: there is a similarly reduced rate for the expanded 25-, 37-, and 67-locus tests from FTDNA as well.) We will need to know something of your male-line ancestry in order to make sense of any DNA matches or near-matches you may have. If your haplotype matches our reconstructed haplotype for Edmund Rice, we will be especially keen to learn more about your Rice/Royce ancestry.

For more information contact our project administrator/coordinator: Bob Rice.
 


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Origin of the Rice Name

The origin of the Rice surname is not known. Indeed, there are clearly many different origins. Click here to see a description of some of the most likely origins, and the relationships among them. It seems likely that the Rice name may blend into other spellings -- not just Royce, but also others, including Reece, Rease, Ris, Ries, and even Price. In other words, the participants of this project need to keep in mind the possibility that their closest genetic relatives may bear a different form of the name. Therefore, if you have no close matches within the Rice DNA project, you may wish to participate in other projects as well, particularly the Reece project and/or the Price project. Similarly, if you are a Reece or a Price with no close DNA matches within your own name, you may wish to join the Rice project.
 


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Contacts

More information about this project: email Bob Rice.

A list of (almost all?) active DNA projects.

For information about the Edmund Rice (1638) Association (and how to join, should you care to do so), please visit our membership page.

There is also a Royce Family Association. You may visit their web page to learn more.

The Reece DNA project has a web page where you may see the test results. Some, but not all, of the members there are also in the Rice project.

The Price DNA project also has a web page.


 


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Results

At the end of this page, we present the DNA test results we have obtained so far. As additional information becomes available, members of the "other" group may be separated out into new groups with identified common ancestors. In cases of ambiguous DNA results, we will depend in part on lineages supplied by the test subjects for determining how the groups should be constructed.

In Tables 1, 2, and 3, each line begins with a unique ID. The 4- and 5-digit ID's refer to FTDNA results. ID's consisting of "N" and a number refer to members of the Genographic Project (tested by FTDNA) who have also joined our project. All other 5-character alpha-numeric ID's refer to entries in YSEARCH. 4-character ID's beginning with the letter "S" refer to Sorenson Genomics/Relative Genetics results. 4-character ID's beginning with "sm" refer to haplotypes found in the Sorenson Molecular Genealogy Foundation Y-DNA database. (See also below under Relevant DNA Projects.)

Nomenclature

The haplotypes are all presented in a consistent nomenclature. For example, pre-2003 FTNDA results for the DYS464 complex have been adjusted by -1, in accord with their change in reporting standards on 2003 May 19. Similarly, pre-2004 results from Relative Genetics have been adjusted by +1 at DYS461, in accord with their change on 2004 June 29. (Our DYS461 result for 1673 came from Relative Genetics in early 2002 and is the only one affected by this particular change.) ID 1673 in Group 1 is a case in point: the same person was tested at three labs (FTDNA, Relative Genetics, and Oxford Ancestors), and the tables include the results from all three. In addition, the pre-2008 SMGF haplotypes have been adjusted by -1 at DYS19, -3 at DYS448, +1 at DYS461, and -1 at Y-GATA H4. The topic of standardization is a thorny one for genealogical DNA testing, since all of the testing companies have changed their reporting standards at one time or another, and they all have disagreements with each other about these standards. (As of 2007 and 2008, FTDNA even disagrees with itself on the standard for Y-GATA H4.)

Data Format

In Table 1, we show only the results for loci included in the FTDNA 25-locus test. Other loci from various sources are shown in Table 2. 30 additional loci from the FTDNA 67-locus test are shown in Table 3. If an entry in the tables is blank, that means the corresponding locus has not been tested. The only exception is the DYS464 complex, which normally has four values reported as a-d, but which may have more or fewer (we have seen 3, 4, 5, and 6 values in members of this project).

The reconstructed ancestral haplotype, if known, of each group is given as the first entry in the group, with the common ancestor's name (if known) as the ID. Individual mutations from the relevant ancestral haplotype are shown in red boldface against a gray background. For loci whose ancestral value is unclear, the background of the whole column is white. We recognize that mutations are inevitable, given enough test subjects and/or the passage of enough time since the progenitor. Nonetheless, these mutations are rare, as the table shows, and the appearance of any discrepancy between the haplotypes of putatively related individuals is cause for concern. (See the discussion of Group 2.) The question is always whether we are so "unlucky" that a rare-but-inevitable event occurred right here, or so "lucky" that a rare coincidence gave two unrelated persons very similar DNA. To resolve that question, we need conventional genealogy (as we do indeed have for the individuals assigned to Group 1).
 


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Test Verification

Subjects 1668, 1669, 1670, 1672, 1673, 4188, 5128, and 5129 all were tested twice independently, once through BYU/Sorenson/Relative Genetics and once through FTDNA. We therefore have cross-checks for 73 of the numbers in Table 1. Unfortunately, there are systematic calibration questions that complicate the comparison for 25 of those numbers. However, 48 of the numbers are straightforwardly comparable, and we find that 46 of the 48 agree between the two labs. This is moderately encouraging, though by no means as good as we expected. In cases of discrepancy, we show the FTDNA result in the table.
 


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Other Relevant DNA Projects

Besides the many other surname-based DNA studies, there are some projects of global scope that include, or may include, Rices and other variants. First of all, there are public databases of user-contributed test results, such as YSEARCH and YBASE . Each of these databases contains test results that members of our project have uploaded. YSEARCH even includes the reconstructed 42-marker haplotype of Edmund Rice. These databases are searchable by haplotype and by surname.

Another database that contains Rices is the SMGF database. This database was initially searchable only by haplotype, but it has allowed surname searches as well since the Fall of 2005. It also has a list of the surnames included and the number of each surname. Initially, there were 4 Rices included in SMGF, and our searches discovered one of them by its haplotype (sm04 in Group 7), in addition to three related men whose family surname is no longer Rice. When the search by surname was implemented, we found the remaining three Rices and one more that has been added since then. To see these data at SMGF, or to look for matches with other entries in Tables 1 and 2, visit our index of SMGF search links.

A fourth database known to contain Rices is that of the Genographic Project conducted under the auspices of the National Geographic Society. This project is not a genealogical one, but is aimed at population studies, such as tracing the paths of human migration over tens of thousands of years. The main thrust of this project is the analysis of isolated indigenous populations around the world, but another component is a study open to the public. Participants in this public study are given their choice of the FTDNA 12-marker Y-DNA test (males only) or the FTDNA mtDNA HVR1 test. In either case, the measured haplotype is used as a predictor of the participant's ancestors' migration routes. After these initial results are returned, the participants are routinely invited to follow up with the genealogical aspects of DNA testing by joining surname studies at FTDNA, and several such persons have joined the Rice project. (Note: the link to the Genographic Project may not work for some web browsers.)

A fifth database of global scope, called the YHRD database, includes anonymous samples from all over the world. This database is aimed at forensic DNA research, rather than genealogy, but it is genealogically useful nonetheless for indicating where a given haplotype can now be found.
 


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Notes and Discussion

Here, we discuss each group of test subjects in turn. One clear message from all of this is that a correct understanding of the relationships requires some conventional genealogy to go with the DNA test results. See especially the discussion for Group 2. You may click on the heading for each group's discussion to view the data for that group in Table 1, just as clicking on the group number within Table 1 brings you back to the discussion in this section.

Group 1

As indicated already, Group 1 consists mainly of male-line descendants of Edmund Rice of Sudbury and Marlborough. This was the first group set up for study and remains the largest. Initially, we focussed on known descendants of Edmund, but, as the project has grown, we have tested more and more subjects who have not been able to trace their lineages back to Colonial times. Some of these have proven to match Group 1 while others have not. In some cases, the conventional genealogical proof of descent from Edmund has been found only after the DNA testing was done. At present, no member of Group 1 has found evidence that he is not descended from Edmund, but that remains a possibility, particularly if and when we find a Group 1 member outside of North America. The following members of Group 1 are still researching their ancestral lines: 6220, 13558, 28544, 39845, 40252, 42343, 86189, 103060, and 110355. Given the new motivation provided by the DNA evidence, it seems likely that some or all of them will eventually find solid conventional links to the rest of the group.

Three members, sm02, sm03, and sm06, were not tested as part of this project, but were instead discovered in the Sorenson Molecular Genealogy Foundation Y-DNA database. These are a father and two sons, and their surname is not Rice, but rather King. Their ancestry goes back to Edmund Rice via Edmund's grandson Samuel King alias Rice. (See the story of the latter's name change in the database of early descendants of Edmund accessible from our main page, where he is listed as "Lt. Samuel Rice King".)

Two other members, 13364 and 24143, have a rather unconventional link in addition to the DNA evidence. In 1704, four Rice boys descended from Edmund were captured by Mohawks at Marlborough (later Westborough), Massachusetts, and carried off to Canada. One was ransomed, but the other three remained and were adopted into the Mohawk tribe. Many years later, one returned to visit Westborough, but he no longer spoke English and had to talk to his relatives through an interpreter. The contact was not maintained, and so there is no collected record of the descendants of these expatriates. Nonetheless, Rice remains a relatively common surname among the Mohawks to this day, and both of these participants are Mohawk Rices. The two match each other 25/25 and match 24/25 with Group 1. 24143 traces back in the male line to one of the Rice captives and thus in turn to Edmund. (However, his lineage was not completed until after we had obtained his results for the first 12 markers. This research was a daunting task.) The fact that the two match each other and share the same surname within the same tribe is compelling evidence that 13364 is closely related and suggests that he may descend from the same captive, but his lineage has not yet been traced all the way. For comparison, we note that a third Mohawk Rice (16023) has also been tested and shows a haplotype very much unlike Edmund's and indeed typical of Amerindian Y DNA, rather than European. Because of the traditional Mohawk system of matrilineal inheritance, it seems likely that anyone in the tribe with the Rice surname may indeed be descended from Edmund, but not necessarily in an all-male line.

A tree diagram of the 25-locus haplotypes in Group 1 shows the majority haplotype at the center and the twelve mutated haplotypes each on a spoke radiating from the center -- a classic "star-shaped" diagram indicative of evolution from a common ancestral type. Not surprisingly, the diagram becomes more complicated with the inclusion of the extended haplotypes shown in Table 2, as 1671 and 1673 share a forked spoke with one mutation in common and one separate mutation each. Interestingly enough, the shared mutation (CDYb=38) can be pinpointed uniquely to Thomas Rice, son of Edmund. This is the first mutation in our project we have been able to tie to a specific individual.

We have upgraded two members of Group 1 to the 67-locus test introduced by FTDNA in the Spring of 2006. These two descend from different sons of Edmund Rice, and thus their perfect agreement on the 30 "new" loci gives us 30 additional loci of the reconstructed ancestral haplotype, for a toal of 72 loci.

Group 2

Group 2 was originally created on the basis of a single sample, mainly because of a lesson it had to teach. However, we have subsequently found other samples that teach the same lesson and have added them to this group, even though there is no evidence at present that any two members of the group are related to each other. Consider the haplotype of sample 3109, the first member of this group. It is sobering to compare this haplotype with that of Edmund Rice. In the first 12 loci (the basic DNA test), these two differ by just one step at one locus, but the additional 13 loci of the expanded test reveal three more differences, including two differences of two steps each. (The contrasting color code in Group 2 shows the loci that differ from Edmund Rice.) Looking only at the basic test, one would be tempted to conclude that the two lines are closely related, but the expanded results show that notion to be false. It is clear now that 3109 is unrelated to Group 1.

The same hazard is lurking for other testees, as we discovered. Therefore, it is often necessary to upgrade to the expanded test when the basic test indicates a possible match. Without the backing of conventional evidence, the DNA results can obviously be misleading. A similar problem (and the same solution) can be seen in the Jarman-German study.

The conclusion we must draw from Group 2 is that sheer coincidence may yield near-matches with the same or similar surnames, based on the existence of a large pool of vaguely similar haplotypes. This vague similarity does, in fact, indicate a relationship among the test subjects, but that relationship apparently goes back thousands of years and falls outside the realm of genealogy. We now discuss the rest of this disparate group.

Interestingly enough, one member of the group (53053) differs by just one step from the first member (3109) at one locus among the first 12 (just as 3109 differs by one step from Group 1), and 53053 goes on to be an exact match with 3109 on the next 13, and now stands just two steps away in all on 37 loci. There is, as yet, no evidence of a genealogical link between them, but both lines have perhaps been traced back to Tennessee, and these two therefore have a reasonable chance of discovering a link.

N1892 is another 11/12 match with Group 1, being only one step away at 12 markers, just as 3109 is. Also, like 3109, he shows three additional differences at 25 markers, including one two-step offset. With only five steps of difference at 25 markers, he falls into the "gray area" for 25-marker comparisons (more than two, but fewer than six steps of difference). However, there is another consideration in his case: he has traced his ancestry to a presumed German immigrant, whose Reiss surname was Americanized to Rice in 1840. Furthermore, he has five additional mismatched markers at 37. It is therefore clear that the seemingly shared surname is really a cross-language coincidence, and the connection probably dates back before surnames were used. N1892 has 26 exact 12-marker matches in the FTDNA customer database, including seven with Finnish email addresses and a like number with American email, but Finnish-sounding names. This, by itself, is far too small a sample to yield any statistical inferences, but the FTDNA ethnic origins database shows that he is an exact 12-marker match with nearly 8% of the samples tested with Finnish ancestry, plus 1% of those with Swedish ancestry and no other country-wide match even approaching 1%.

22348's results are only five steps away from some of the members of Group 1 at 25 markers (and only two steps at 12 markers). As such, he falls (barely) into the "gray area" with respect to those individuals. (For 12-marker comparisons, the "gray area" is more than one, but fewer than four steps of difference.) However, comparing a modern individual and a reconstructed group progenitor is much more stringent than comparing two modern individuals -- there are two separate lineages where mutations could accumulate in the latter case, but only one lineage in the former. Thus, we can be quite confident that 22348 is not a descendant of Edmund Rice, just as in the cases of the other members of Group 2.

A tree diagram of Group 2 would be a hopeless snarl, with duplicated parallel branches criscrossing each other and leaving many gaps unoccupied by actual samples. Such a pattern indicates a very much older and larger population that is only sparsely sampled. This point re-emphasizes that the members of this group are not related within a genealogical time frame.

Group 3

The members of Group 3 are identified as male-line descendants of Robert Royce (c1606-1676) of New London, CT. Five of these have been traced back to Robert Royce with reasonable certainty by conventional means, seven others tentatively, seven are not yet documented, and three others were previously thought to be descendants of Edmund Rice. However, the near-exact match of all the samples that have been tested on 25 loci makes it seem likely that at least these are closely related, and probably all of them. Since the testees with documented genealogies back to Robert Royce agree at least 24/25, and include exact-match descendants from two different sons of Robert, the group as a whole is now quite firmly established, even though the other members have not traced their lineage back to the founder. We have therefore reconstructed the ancestral Royce haplotype and included it in the table. In any case, this group shows the same sort of "star-shaped" tree diagram as Group 1: a central consensus haplotype with several adjacent ones showing one or two members each.

Three members of Group 3 have been extended to 37 loci, including two members who trace back to two different sons of Robert Royce and a third member who has tentatively traced back to yet another son. All three agree exactly on the last 12 loci.

Two members of this group, a father and son, submitted their samples and pedigrees to the SMGF project, where we found them and designated them sm18 and sm19. We have not yet verified their lineage from Robert Royce (and we therefore consider them "tentative" genealogically). They are also tentative genetically because they have four differences from the Group 3 haplotype out of the first 25 markers. Nonetheless, they match on all the rest of the markers (18 more in all), and it seems most likely that the four differences are simply a statistical fluke.

More recently, a participant (100323) has been found who matches them on all but one marker among the first 25, including three of their four mutations. Two of these three mutations are shared with one or more other members of the group, but not both at the same time. In any case, 100323 is closely related to sm18 and sm19 according to the stated lineages, and the fact that he shares only three of the four mutations with them improves the fit of all of them into Group 3.

Shared mutations

Just as in Group 1, where 1673 and 3111 share both a common line of descent and a mutation from the ancestral haplotype, we see three shared mutations in Group 3: DYS19=15->16, DYS385b=16->17, and DYS393=14->13. In principle, the established lineage of one member of such a cluster can serve as a useful clue for any others lacking a clear lineage.

However, three of the four in the first cluster have traced back to Robert Royce with no intermediate common ancestors, and therefore this clue is contradictory for the remaining member. Of course, such clues do not always bear fruit, even without contradictory indications, since duplicate mutations can arise independently, unlikely though that is.

The second cluster led to a search for the exact timing of the mutation. In the case of 12451 and 15824 (and 15824's near kin 21622, 33906, and 33908), we now have evidence that the mutation at DYS393 did indeed occur twice independently, since 15824's fourth cousin 21032 (and third cousin 33907) lack the mutation. By recruiting all of these cousins for testing, 15824 has now demonstrated that the DYS393=14->13 mutation occurred in his great grandfather independently of the duplicate mutation seen in 12451.

The third cluster is also problematic, since two of the testees belong to two such clusters, thus assuring that at least one mutation has been repeated independently.

The second cluster is of interest for another reason. It includes a striking example of an unusual process that has only recently been found in the Y chromosome -- known variously as "gene conversion" or "recombinant loss of heterozygosity" or "doubling." Comparison of the results for 15824 and his great-grandson 33906 shows a change on DYS459a,b from 8,10 to 10,10 as well as a change on DYS464a,b,c,d from 11,14,14,15 to 14,14,14,14 and on CDYa,b from 33,38 to 38,38. We see an apparently simultaneous replacement of an "8" by its partner "10" and of "11,15" by their partners "14,14" and of "33" by its partner "38." Since all three of these markers occur on the same palindrome of the Y chromosome, it seems virtually certain that one of two possible events has taken place between the two individuals: (A) a large-scale replacement, such that a long segment on one arm of the palindrome has been converted into a copy of the corresponding segment on the other arm, or (B) the long segment was simply lost. Unfortunately, the lab tests are not well enough calibrated to detect the difference between duplicate markers and missing markers, and the lab reports always indicate duplicates in such cases. We are therefore unable to choose between the two possibilities, despite the appearance of certainty in the lab reports.

Group 4

Group 4 was created based on the exact matches at 12 loci for three samples, now grown to sixteen, plus four further samples that match the others 11/12. Twelve have been extended to 25 loci, and four of these match each other exactly 25/25, and two other match each other as well. Indeed, if we ignore the DYS464 complex, we have seven 21/21 matches, but there is a complication at DYS464: although all twelve have two 15's, and eleven have at least one 17 among the four copies of this locus, there is no consensus on the fourth copy. Seven have another 17, while four have a 14 and two have a 16. It seems likely that the 14-vs-17 discrepancy is due to a single mutation, but it is uncertain which is the ancestral value. 17 is favored by the fact that it is most common, but it is easier to postulate a 14 mutating into 17 in this family than a 17 to 14. We could resolve this question if we had clear lineages for all twelve testees, but the relationships have not been tied down. We have therefore left DYS464c blank in the reconstructed ancestral haplotype for this group.

In any case, it seems likely that the 14-17 mutation in DYS464 occurred relatively early and that it therefore splits the group into two subgroups which could be distinguished genealogically. It would therefore be useful for the rest of the members to upgrade to 25 to help in the sorting of the two subgroups. In fact, we have already used this splitting to characterize the discrepancies we see in Table 2 for the four members of this group who have upgraded to 37 markers. Two of the discrepancies appear to be individual line mutations (at DYS576 and DYS570), but one (CDYa) appears to split the group in the same way as DYS464.

Although nine of the donors have tentatively been traced back to Rices of Virginia, and eight others to Rices of Kentucky or the Carolinas (and probably to Virginia ultimately), the identification of the progenitor remains uncertain. Indeed, some other testees who do not match seem to be contending for the same progenitor (Thomas Rice of Gloucester Co, Va., c1650 - c1716). It will be necessary to test more descendants to firm up this group.

One apparent member of the group is sm09, discovered in the SMGF database. He is in the odd position of differing on three markers among the first 12 but agreeing on all the rest, being 29/34 with 4131 and 56745 and 28/34 with 17939. sm09 has been traced back to 1825 in North Carolina and thus fits in geographically with the group but cannot yet be tied in genealogically.

Four members of Group 4 have upgraded to the 67-locus test introduced by FTDNA in the Spring of 2006, and their results agree perfectly for the 30 "new" loci. This agreement reaffirms the consistency of the group as a whole. However, since the relationship among these four members is not known, we have not used them as a basis for predicting the ancestral values of the new loci.

Group 5

This second Virginia Rice group is a similar collection of shared and almost-shared ancestry, bound together by the matching DNA. Until recently, the arguable common ancestor of the whole group was one William Rice born c1756, thought to be the father of three men known to be the ancestors of five of the members of this group. No other documented ancestors of the group were early enough to challenge him. Now, however, other lines have been discovered going back to contemporaries of William who may have been his brothers. If so, the new candidate common ancestor would be their father, possibly a certain James Rice who died in Ohio in 1844 at a very advanced age (said to be 120 years old!).

In the process of constructing this group, the DNA evidence was at first inconclusive, since there was one discrepancy among the first 12 loci. However, extending the tests revealed a perfect match among the 13 additional loci for the two subgroups. This overall 24/25 match is a fairly convincing demonstration of relatedness.

13040 has been traced only as far back as 1845, but matches the main haplotype 12/12. 17265 also matches but has not been traced all the way back. We therefore include them in this group pending further research.

Note: we are assuming that the three subjects 5894, 6838, and 48417 represent a side branch of this group, since their lineages have not been traced, as most of the others have. Therefore, we show the discrepancies as mutations in these three haplotypes. However, given the small number of testees in the group, we have to entertain the possibility that these three are the "main" branch, with the rest being a "side" branch. This issue would be resolved if we knew how the two subgroups are related.

Additional testing could also help clarify the picture. Eventually, as Rices of unknown lineage are randomly added to the project, the most common haplotype in Group 5 is more and more likely to be the ancestral, "main" branch. The originally discovered haplotype in Group 5 is still the most common, but the selection of test subjects has not been predominantly random, and so that line of argument is only theoretical.

Group 6

This third Virginia group has only three representatives so far and includes only 12-locus tests, but it seems to be well documented all the way.

Group 7

This fourth southern Rice group has been designated on the strength of its growing numbers. Reports of the earliest known ancestors range from Tennessee and Kentucky to North Carolina. Most members of this group match each other 23/25 to 25/25. These test subjects thus appear to constitute another group, even though there is as yet no agreement on a common ancestor.

Originally, three other testees were tentatively included in Group 7, but extending the tests to 37 markers has shown that the relationship is too distant to fit all into one group. (See Group 7A.)

Based only on the first two panels of markers (as shown in Table 1), subjects 44461 and sm04 appear to partly bridge the gap between Groups 7 and 7A, since they share a mutation at DYS455 with 7A and lack a mutation at DYS439. 44461 also shares DYS464c=16 with 7A, while sm04 does not. Both of them have the normal four copies of DYS464, unlike the members of 7A. Moreover, the extended results shown in Table 2 give a different picture. 44461 matches 12/12 on the third panel with 5820 (the only other member of Group 7 to test all 37) and 7/7 with sm04. Thus, 44461 is definitely not a bridge between the groups. sm04 is slightly less clear. Of the seven extended loci in common between FTDNA and the SMGF, sm04 matches these two extended Group 7 members perfectly, but only 5/7 or possibly 6/7 with Group 7A. However, he has no test results for the other five extended loci, where four of the discrepancies between 7 and 7A are found, and it is therefore difficult to place sm04 with complete assurance. sm04 traces his line to William Littleton Rice, born 1836 in Arkansas, and probably further to Tennessee and Virginia. However, he remains anonymous, and the details have not been verified. Another possible route to the proper placing of sm04 (and further investigation of the relationship between the groups) would be to obtain test results from SMGF on some of the other members of Groups 7 and 7A.

Group 7A

This group was originally thought to be part of Group 7, but further testing has shown the two groups to be separate.

5796 was the first member of this group. He differs from Group 7 by two ordinary one-step mutations among the first 21 markers, some anomalous results for the DYS464 markers (he has only three copies instead of the usual four), and six or seven one- or two-step mutations in the third panel. The second member to be added was 6977, a third cousin once removed of 5796. 6977 has five copies of DYS464, but otherwise matches 5796.

Test subject 37139 also appears to be closely related to 5796. He shares both of the distinguishing mutations (DYS439=13 and DYS454=11) that set 5796 and 6977 apart on the first 21 markers, and he also has an unusual number of copies of DYS464. Oddly enough, despite having an extra copy of DYS464, just like 6977, he is two steps away on those five markers. Still, the mere fact of having an extra copy of DYS464 is very suggestive, and the comparison between 37139 and 5796 shows 31/33 agreement on the remaining loci.

Group 8

This group descends from the Rices of Weymouth, MA, who are now confirmed to be unrelated to all the other known Rice/Royce lines. Note: it was a member of this family who founded Rice University.

There is now a 23/25 near-match with another Rice, 78290, who has so far found no indication of a connection with the Weymouth line. Such a match is neither close enough nor distant enough to give an unequivocal answer to the question of whether 78290 is paternally related to Group 8 in a meaningful time frame. Perhaps, if one or more members of the group were extended to a 37-marker test, the expanded comparison would give a clearer answer, one way or the other.

Group 9

Although testee 14746 was initially placed in Group 5 based on a 12-marker comparison, the 25-marker results showed him to be quite remote from that group. Since then, the 12-marker results for nine more test subjects have matched 14746 exactly. There are now so many that they appear to form a separate group. Of course, we cannot assign permanent membership without convincing 25-marker tests or documentary evidence, but we moved these results into a tentative group by themselves in the expectation that further testing would confirm the relationship. Nine of the ten have now been extended to 25 markers, and the results are mostly convincing. Two subgroups of three haplotypes each match among themselves exactly. Four haplotypes differ from what seems to be the ancestral pattern only at one locus in DYS464, one being an ordinary mutation of one step, two being offsets of two steps probably due to recombination, and one being apparently a three-step mutation. The latter could also be interpreted as three separate one-step mutations, but such a compound mutation is very unlikely, even more so than a single three-step. These eight subjects match each other far more closely at 25 markers than they do anyone else in the project, and their differences are consistent with a common ancestor within a genealogical time frame.

The remaining 25-marker haplotype, 36743, differs by two steps from the consensus, and both of these differences match the members of Group 5. Thus, although 36743 is closer to Group 9 than to any other group, it turns out that Group 5 is a close second. This participant is also notable for bearing the surname Reece, instead of Rice. As noted above, a surname variation at this level should not be taken too seriously, but it does stand as an additional factor in the group assignment. Thus, this haplotype's membership in the group remains a bit more tentative than the others. Finding a conventional genealogical link or a point where Reece changed to Rice, or Rice to Reece, would help to solidify the connection, but it remains possible that the link is an ancient one dating back before the adoption of surnames.

One other member has been tentatively added to this group on the basis of an exact 12/12 match. Again, like all the rest of Group 9, this one is only one step away from some members of Group 5, and this assignment needs a 25-marker comparison for confirmation.

Group 10

Although the two members of this group match each other only 24/25, they have family lore saying that their great great great grandfathers were brothers.

Group 11

This group has a perfect 25/25 match and genealogies pointing to a common ancestor in Tennessee.

Group 12

This group, like Group 11 above, also has a perfect 25/25 match and genealogies pointing to a (different) common ancestor in Tennessee. This was the first real group to spring from the cluster of unmatched, but similar, haplotypes we designate Group 2. As such, the members have some near-matches in Group 2, and even in other real groups, as well. For example, there is a member of Group 17 who is only three steps away from Group 12 in a 25-marker comparison. They could conceivably be related, since a difference of three steps falls in what we would normally call the "gray area". However, such a relationship is not suggested by conventional research (nor helped by the fact that the other member of Group 17 is four steps away). Again, the fact that so many haplotypes lurk in the vicinity of Group 2 means that the standard of proof must be higher for designating groups among them.

Group 13

This is yet another group with roots in Virginia. The progenitor, Samuel R. Rice, was born there in 1744, believed to be the son of another Samuel. Descendants of two of his sons have been tested, and three of the descendants have results for 25 loci. From these results, we can reconstruct the ancestral haplotype for 24 of the 25 loci, but, unfortunately, one locus is uncertain. All three descendants of the eldest son agree with each other on that locus, and the one descendant of the younger son differs. Although it is somewhat more likely that the three in agreement represent the ancestral value for that locus, we cannot be sure until we find the same value in a descendant of another son.

Group 14

Like Group 12, this group has a DNA pattern very similar to that of Group 1, and the first two members of Group 14 were initially placed in the catch-all "Group 2" until a third match was found. The members' relationship to each other is based on tracing their lines back to Pennsylvania. The documentation is still lacking to connect them all to one common ancestor, but the 25/25 match among them definitely supports the connection. Even so, this group was not deemed "solid" until there were three members, because of the other (apparently unrelated) members of Group 2 who were also genetically similar.

Test subject 14295 (the first member of this group) has no fewer than three near-matches in the "gray area" of 12-marker comparisons, as well as one 11/12 match, in addition to his matches with the rest of Group 14. Needless to say, the others in the group share the same near-matches.

First, they differ from Group 1 at two loci, by one and two steps, respectively. Even factoring in the possibility that the two-step difference stems from a two-step mutation, the most likely time of the common ancestor is about 14 centuries ago, long before the Rice surname was adopted. Extending the test to 25 loci revealed one further two-step difference (at DYS448, where all members of Groups 2, 12, and 14 differ from Group 1). The most likely estimate of the common ancestor, based on a 25-marker comparison, is about 10 centuries, though a separation of only four centuries is still within the realm of possibility.

Second, they differ from 7648 in Group 17 at two loci, again by one and two steps. The 12-marker-based common-ancestor estimates are the same for this case, but the 25-marker comparison gives an estimate of about nine centuries.

Third, they differ from Group 6 by only one step at each of three loci. The 12-marker common-ancestor estimates for Group 14 and Group 6 are similar to the above, but a century or so more remote. Indeed, since no member of Group 6 has been extended to 25 loci, we would need several extensions in order to probe this possible link.

Finally, they differ from 17633 (in Group 2) by only one step on 12 markers, or three steps on 25 markers. Indeed, one member of the group has extended to 37 markers, as has 17633, and the difference there is six steps. These distances all fall into the "gray area". In any case, there is (as yet) no conventional evidence pointing to a link. Without such evidence, we have no grounds to suppose a link exists within genealogical time, even though such a link is possible.

Group 15

The members of this group have a haplotype that belongs to a cluster commonly found in northwestern Ireland, but there is no conventional evidence pointing beyond the immediate origins of this group in Rhode Island. Indeed, they do not yet have a documented link between them, but their 36/37 match, coupled with the fact that both 4507 and 86858 trace their lines back to Rhode Island, provides convincing evidence of a link.

There are several other members of the project who come close to this group, but not close enough to be convincing and not backed up by any conventional evidence.

Group 16

The members of this group are a very close match (36/37) and have a genealogical connection as well, albeit a connection based in part on undocumented sources. They also have a very close match (36/37 for one of them) with a group of Fletchers from the same area of Virginia. It is not known what connection there may be between these Rices and Fletchers. Since the genetic match is not perfect, the separation between the two families could date back as much as five centuries, i.e., long before the earliest known ancestors of either group, though the odds favor a more recent connection.

Group 17

This is yet another group that is close to the pool of similar, but evidently unrelated, haplotypes in Group 2. Indeed, subject 7648 makes an interesting case study. He was one of the fairly early participants in the project, and he came close genetically to many others along the way, but it was five years before a real match came along. The long delay was especially surprising because his male-line ancestry had been traced back to early Colonial Massachusetts, and so there should (in principle) be many of his lineage to be tested. Suffice it to say that a match eventually did appear. The common ancestor of Group 17, Samuel Rice, is believed to be the son of Massachusetts immigrant John Rice of Dedham, who married there in 1649. Since John Rice had another son who raised a family, there is a good possibility that we can extend the group back one generation by testing a descendant of the other son. This would have the added benefit of producing the first real evidence of a connection between John and Samuel (beyond the mere fact of living in the same town). Let us hope it won't take five more years!

This haplotype falls just short of matching Group 1 and also another subject (5204) currently in Group 2. In the first 12 markers, 7648 differs by just two steps from Group 1 and by three from 5204, roughly splitting the difference between the two extremes. Such a result is difficult to assess, since it falls into the "gray area". A 12-marker difference of three steps is ordinarily enough to show two subjects are unrelated on a genealogical time scale, and even two steps of difference would be very unusual within the same family. However, a large enough population will inevitably include a few outliers, and the best way to detect such outliers as such is to test more markers. We therefore encouraged 7648 to extend from 12 to 25 markers. With this broader base, we found that 7648 differs by five steps (including a two-step difference on one marker) from Group 1 and by six steps from 5204, thus giving a much more statistically secure indication that no close relationship exists among them. Two more close-but-not-close-enough matches for 7648 also came to light. First was a near-match with 21119 and 41527 in Group 12, manifested as a one-step difference at 12 markers. This is the same level of difference as seen at 12 markers between 3109 and Group 1, and the same warning applies here. Without explicit conventional research and/or additional DNA testing, the 12-marker near-match is merely a clue, and possibly a misleading one. Extending the comparison to 25 markers revealed a total of three one-step differences. This result was still in the "gray area," and we could neither rule out a connection by the DNA comparison, nor find any evidence of a connection by conventional means. The other near-match was two-step, 10/12 with 21748. Extending this comparison to 25 markers showed a difference of 8 steps in all -- no longer of any interest.

When the real match finally appeared, it was 12/12 and 24/25. Just as importantly, though, was the fact that 111320 had also been traced back to the same immigrant Rice. This concurrence of the two types of evidence makes a convincing case that both types are correct.

Group 99

In the "other" group, we are finding considerable diversity, with at least 41 more distinct haplotypes identified already, and only four matches among them. One match, an identical match between 4090 and 5070, is no surprise at all because those two are brothers. The second match is 11/12 between 3145 and 21802, but extends to 21/25 and thus falls into the "gray area". 21802 has now extended to 37, and, if 3145 does likewise, we might be able to get a clearer determination of the relationship.

The third match is a five-way match at 12 markers, with two identical pairs one step apart and another member one step from one of the pairs. The bridging pair, 87439 and 100521, are both tested to 25 markers, and they match each other on 23 of the 25, with three steps of difference in all (thus putting them in the "gray area" on the available tests). Indeed, other participants are nearly as close on 25 markers to one or the other of these two as the two of them are to each other. Moreover, these five are very close to the most common DNA type in western Europe, and it is necessary to extend all of them at least to 25 markers before the match could become convincing evidence of a link. The potential for a near relationship is there, but we would need some conventional evidence to confirm that possibility.

The fourth match is three-way at 12 markers, with two exactly the same (35599 and 88027) and another (9166) one step away. The non-exact member and one of the pair have upgraded to 25 markers, and these two have three discrepancies in all of one step each, thus putting them in the "gray area". The two exact matches share not only their 12-marker haplotype, but also the Reece variant of the surname, and they would do well to compare at 25, since they may find convincing evidence of a recent link.

There was briefly just a hint of a possible larger grouping of subject 4283 with what subsequently became Group 7 and Group 16. At the time, only one member of each of those groups had been found, and those two, along with 4283, appeared to fall into the "gray area" between one and four steps of difference. In that case, there was no pair as close together as two steps apart, but it was still intriguingly close. Here again, we relied upon an expanded test to cast further light on the situation. The three candidates were all extended, and they showed large differences between every pair of them. Thus, the suggestion of a connection fizzled here as well. Nonetheless, we subsequently found matches for two of the three and thus established groups 7 and 16.

There is yet another hint of a possible distant connection between subjects 4798 and 6510, who differ by one step at each of three loci in the 12-marker test. 4798 has now been extended to 25. If 6510 does likewise, and they still have only the three differences, or if intermediate subjects come to light who bridge the gap between these two, and if conventional evidence of a link could be found, then we would probably have a new group identified. Meanwhile, we are counting these two as distinct. Indeed, 4798 had a second possible connection, being only two steps away from 25686 on the 12-marker test. Again, the 25-marker gave a more precise reading; the two steps of difference turned into five steps, still within the "gray area," but just barely. Without specific evidence of a relationship, such a wide gap is probably not worth pursuing.

During most of the year 2003, subject 7897 was tentatively assigned to Group 7 on the basis of a 12-marker test, even though his 12-marker haplotype differs by two steps from the consensus of that group. The reason for this assignment was the presence of two other tentative members of the group who shared one of the two apparent mutations in 7897. Now, however, the 25-marker results show that 7897 is not closely related to anyone in Group 7, and he has been moved to Group 99. Similarly, another test subject, 59548, was tentatively placed in Group 7A based on the first 12 markers, but has now been moved to Group 99 because the extension to 25 shows no similarity. Yet another test subject, 14746, was initially assigned to Group 5 based on the same type of similarity to "atypical" members of that group, and the extension to 25 markers had the same outcome: 14746 was also moved to Group 99. However, others have now been found who match him, and these matches have now been moved, yet again, to a new group by themselves (Group 9).

17791 has a possible (but not probable) connection to Group 15. Of the first 12 markers, he matches on 11. However, their extended test results match only 21/25, thus falling into the 25-marker "gray area" (more than two, but fewer than six steps of difference). By the same token, 36732 has a 12/12 match with Group 15, but here the 25-marker comparison is even worse: six steps (same as the 25-marker comparison between 17791 and 36732). These test subjects are all at or near the most common haplotype of northwestern Ireland, and there are therefore many apparently close haplotypes with only a distant genetic connection. Thus, it is necessary in these cases to compare at least 25, and perhaps 37, markers to get convincing evidence of a recent connection, unless some conventional genealogical evidence can be found.

A haplotype that comes moderately close to Group 7A is not fully filled in. sm27, found in the SMGF database, has only 21 of the first 25 markers. If the omission of DYS464 signifies that sm27 has five or more copies of this marker, then he may indeed belong to 7A. Until we have DYS464, we cannot place him properly, and we are leaving him in Group 99 for the time being.

One member of this group (qqjdr) tested independently at Relative Genetics and reported the results to us. We have not found any matches within our project and thus include the results in Group 99.

In the data tables, the haplotypes are arranged in numerical order, except that those with possible matches are grouped together and shaded a slightly darker gray, and the columns of the discrepant markers are displayed on a white background.

Mutation Rate

This study, although too small to provide a statistically significant measurement of the mutation rate in Y-DNA STR markers, does at least provide an estimate of that poorly-known parameter. We have expanded the mutation study to include all confirmed lineages in Groups 1, 3, 5, and 6 as of 2004. Samples from Group 1 (1670-3, 4188, 5128-9), Group 3 (3156, 6061, 7242, 7628), and Group 5 (3869, 4765, 4808, 5022, 8850) represent 125 separate father-to-son transmission events with 25 loci measured; Group 1 (3111-3113, 5300, 6336) and Group 6 (4046, 4641, 5032) have 44 more with 12 loci; and S002-S009 have 40 more with 9 loci. There are thus 125x25 + 44x12 + 40x9 = 4013 mutation opportunities. The mutation seen in both 1673 and 3111 is undoubtedly one and the same mutation, since these two men share several generations of common ancestry. (However, note for comparison that the same mutation has also occurred independently at least twice in Group 1 - in 19847 and 87949. The fact that the marker in question is DYS439, which has a relatively high mutation rate, increases the chances of parallel mutations.) The mutation in 5129 is an astonishing three steps. There is no way to be sure whether this is one mutation of three steps all at once or three separate events of just one step each that coincidentally happened to fall on the same locus and in the same sense, but the latter possibility is so unlikely (five-in-a-million) as to be scarcely worth considering. In contrast, the chance of a 3-step mutation somewhere in this line is probably on the order of one in a thousand. Pending further investigation, then, we will treat this as a single event. Thus, we derive a mutation rate of 6/4013 = 0.0015, with a broad uncertainty.

Note: it may eventually be possible to include the results from other groups in this analysis of the mutation rate, but their ancestral haplotypes are still too uncertain at present.

A combination of many such studies based on deep-rooted pedigrees could yield a significantly more precise value. Preliminary results from a combination of 13 studies shows an average rate of 0.0023 mutation per generation per marker, with a statistical standard deviation of 0.0003. Ultimately, of course, each locus has its own individual mutation rate, which should be taken into account, but the average over all loci is the really important statistic. Indeed, for our purposes the only question that matters is: Does the likely separation allow for a connection in the time since surnames were adopted, or not?
 


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DNA Data

Table 1. Rice Haplotypes: measured marker lengths.
(click on an arrow at the end of a row to see the continuation in Table 2)
Note: DYS464 has been converted to new nomenclature as of 2003 May.
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 1 (Edmund Rice, c1594 ENG - 1663 MA)
Edmund 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
1670 13 24 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
1671 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
1672 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
1673 13 23 14 10 14 14 11 14 12 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
3111 13 23 14 10 14 14 11 14 12 12 11 28
3112 13 23 14 10 14 14 11 14 11 12 11 28
3113 13 23 14 10 14 14 11 14 11 12 11 28
4188 14 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
5128 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
5129 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 20 16 18 28 12 14 15 16 >
5300 13 23 14 10 14 14 11 14 11 12 11 28
6220 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
6336 13 23 14 10 14 14 11 14 11 12 11 28
11268 13 23 13 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
13364 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 14 16
13558 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 27 12 14 15 16
14979 13 23 14 10 14 14 11 14 11 12 11 28
17599 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
19847 13 23 14 10 14 14 11 14 12 12 11 28
20401 13 23 14 10 14 14 11 14 11 12 11 28
22912 13 23 14 10 14 14 11 14 11 12 11 28
23423 13 23 14 10 14 14 11 14 11 12 11 28
24143 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 14 16 >
24992 13 23 14 10 14 14 11 14 11 12 11 28
28544 13 23 14 10 14 14 11 14 11 12 11 28
28661 13 23 14 10 14 14 11 14 11 12 11 28
29880 13 23 14 10 14 14 11 14 11 12 11 28
39845 13 23 14 10 14 14 11 14 11 12 11 28
40252 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
42343 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
63613 13 23 14 10 14 14 11 14 11 12 11 28
86189 13 23 14 10 14 14 11 15 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
87949 13 23 14 10 14 14 11 14 12 12 11 28 15 8 9 8 11 23 16 18 26 12 14 15 16
96421 13 23 14 10 14 14 11 14 12 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16
103060 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 20 16 18 28 12 14 15 16
110355 13 23 14 10 14 14 11 14 11 13 11 29 15 8 9 8 11 23 16 18 28 12 14 15 16 >
sm02 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
sm03 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
sm06 13 23 14 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 18 28 12 14 15 16 >
sm30 13 23 14 10 14 14 11 14 11 12 28 15 8 9 8 11 16 18 28 12 14 15 16 >
S002 13 23 14 10 14 14 14 12 11 >
S005 13 23 14 10 14 14 14 12 11 >
S006 13 23 14 10 14 14 14 12 11 >
S007 13 23 14 10 14 14 14 12 11 >
S009 13 23 14 10 14 14 14 12 11 >
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 2 (Not related, but similar to Group 1)
3109 13 23 15 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 20 29 12 14 14 15 >
53053 13 22 15 10 14 14 11 14 11 12 11 28 15 8 9 8 11 23 16 20 29 12 14 14 15 >
5204 13 22 14 10 13 15 11 14 11 13 11 30 15 8 9 8 11 23 16 20 29 12 15 15 16
17633 13 22 14 10 14 15 11 16 11 12 11 28 16 8 9 8 11 23 16 20 29 12 14 15 16 >
21748 13 22 15 10 13 14 11 14 11 12 11 29 14 8 9 8 11 23 15 21 30 12 14 15 16
22348 13 23 14 11 14 14 11 14 11 11 11 27 15 8 9 8 11 23 16 20 28 12 13 15 16
34698 13 22 15 10 13 14 11 14 11 12 11 28 15 8 9 8 11 23 17 20 28 12 14 15 15 >
102098 14 22 14 10 13 14 11 14 11 12 11 28 15 9 9 8 11 22 16 20 28 14 14 15 15
N1892 13 23 14 10 14 14 11 14 10 12 11 28 15 8 9 8 11 23 16 20 28 13 14 15 15 >
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 3 (Robert Royce, c1606 Som, ENG - 1676 CT)
Robert 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
1668 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 16 >
1669 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
3156 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
3730 14 23 15 10 15 17 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
3758 14 23 16 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
5333 14 23 15 10 15 16 11 13 11 14 12 32
6061 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
7242 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
7360 14 23 15 11 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
7628 14 23 16 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
9941 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
12451 13 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
19375 14 23 15 10 15 16 11 13 11 14 12 32
21032 14 23 15 10 15 16 11 13 11 14 12 32
25714 14 23 15 10 15 16 11 13 11 14 12 32
15824 13 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
21622 13 23 15 10 15 16 11 13 11 14 12 32
33906 13 23 15 10 15 16 11 13 11 14 12 32 15 10 10 11 11 25 14 20 26 14 14 14 14 >
33907 14 23 15 10 15 16 11 13 11 14 12 32
33908 13 23 15 10 15 16 11 13 11 14 12 32
39486 13 23 15 10 15 16 11 13 11 14 12 32
44786 13 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
47510 14 23 15 10 15 16 11 13 11 14 12 32
88798 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15
90330 14 23 16 10 15 16 11 13 11 14 12 32
100323 14 23 16 10 15 17 11 13 11 14 12 32 16 8 10 11 11 25 14 20 26 11 14 14 15
N31121 14 23 15 10 15 16 11 13 11 14 12 32
9uxtm 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
sm18 14 23 16 10 15 17 11 13 11 14 12 32 16 8 10 11 11 26 14 20 26 11 14 14 15 >
sm19 14 23 16 10 15 17 11 13 11 14 12 32 16 8 10 11 11 26 14 20 26 >
sm31 14 23 15 10 15 16 11 13 11 14 12 32 15 8 10 11 11 25 14 20 26 11 14 14 15 >
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 4 (Rice of Virginia)
? 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17
3110 14 24 14 11 12 15 12 12 12 12 13 28
4131 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17 17 >
4086 14 24 14 11 12 15 12 12 12 12 13 28 19 9 9 11 11 25 15 19 31 15 15 16 17
4897 14 24 14 10 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17 17
5076 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17 17
5332 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 14 15 15 17
17939 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 14 15 15 17 >
6276 14 24 14 11 12 15 12 13 12 12 13 28
18407 14 24 14 11 12 15 12 12 12 12 13 28
19119 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 14 15 15 16 >
20488 14 24 14 11 12 15 12 12 12 12 13 28
20980 14 24 14 10 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 32 15 15 17 17
37250 14 24 14 11 12 15 12 12 12 12 13 28
45813 14 24 14 11 12 15 12 12 12 12 13 28
46459 14 24 14 11 12 15 12 12 12 12 13 28 19 9 9 11 11 25 15 19 31 15 15 17 17
56745 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17 17 >
57438 14 24 14 11 12 15 12 12 12 12 13 28
64414 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 31 15 15 17 17
90008 14 24 14 11 12 15 12 12 12 12 13 28 18 9 9 11 11 25 15 19 30 14 15 15 17
N4403 14 24 14 11 12 15 13 12 12 12 13 28
sm09 14 23 14 11 12 15 12 12 13 12 13 29 18 9 9 11 11 25 15 19 31 15 15 17 17 >
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 5 (Rice of Virginia II)
? 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
3869 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
4765 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17 >
4808 13 25 14 10 11 16 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
5022 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
5894 13 25 14 11 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
6838 13 25 14 11 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 31 15 15 16 17
8850 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
13040 13 25 14 10 11 15 12 12 12 14 13 30
17265 13 25 14 10 11 15 12 12 12 14 13 30
21583 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
48014 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
48417 13 25 14 11 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
70565 13 25 14 10 11 15 12 12 12 14 13 30 16 9 10 11 11 25 15 19 30 15 15 16 17
N2333 13 25 14 10 11 15 12 12 12 14 13 30
N46489 13 25 14 10 11 15 12 12 12 14 13 30
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
4
6
4
e
ID
Group 6 (James Rice, c1740 - 1817 Loudon Co, VA)
James 14 22 14 10 13 13 11 16 11 12 11 28
4046 14 22 14 10 13 13 11 16 11 12 11 28
4631 14 22 14 10 13 13 11 16 11 12 11 28
5032 14 22 14 10 13 13 11 16 11 12 11 28
Group 7 (Rice of mid-South)
? 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 30 15 15 17
4091 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 30 15 15 17 17
4165 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 29 15 15 15 17
4462 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 30 15 15 17 17
5820 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 30 15 15 15 17 >
8232 13 24 14 11 12 14 12 12 12 13 13 29
19956 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 12 25 15 19 30 15 15 17 17
44461 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 11 25 15 19 30 15 15 16 17 >
55145 13 24 14 11 12 14 12 12 12 13 13 29
56852 13 24 14 11 12 14 12 12 12 13 13 29
66101 13 24 14 11 12 14 12 12 12 13 13 29
sm04 13 24 14 11 12 14 12 12 12 13 13 29 17 9 10 11 11 25 15 19 30 15 15 17 17 >
Group 7A (Rice of mid-South II)
5796 13 24 14 11 12 14 12 12 13 13 13 29 17 9 10 11 11 25 15 19 30 15 - 16 17 >
6977 13 24 14 11 12 14 12 12 13 13 13 29 17 9 10 11 11 25 15 19 30 15 15 16 17 17
37139 13 24 14 10 12 14 12 12 13 13 13 29 17 9 10 11 11 25 15 19 30 14 15 16 17 18 >
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
4
6
4
e
ID
Group 8 (David Rice of Weymouth, MA)
David 13 23 14 11 11 14 12 12 12 12 14 28 17 9 10 11 11 24 15 19 30 15 15 16 18
17960 13 23 14 11 11 14 12 12 12 12 14 28 17 9 10 11 11 24 15 19 30 15 15 16 18
20221 13 23 14 11 11 14 12 12 12 12 14 28 17 9 10 11 11 24 15 19 30 15 15 16 18
21597 13 23 14 11 11 14 12 12 12 12 14 28
Group 9 (Rice of Virginia III)
14746 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 14 15 16 16
20645 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 16 16 17
24104 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 16 16 18
25156 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 15 16 16
26317 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 15 16 16
27358 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 16 16 17
30512 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 16 16 17
36743 13 25 14 11 11 15 12 12 13 14 13 30 17 9 10 11 11 25 15 19 29 15 15 16 17
89613 13 25 14 11 11 15 12 12 13 14 13 30 17 9 11 11 11 25 15 19 29 15 15 16 16
N35005 13 25 14 11 11 15 12 12 13 14 13 30
Group 10 (Rice of western Virginia)
29562 13 24 14 11 11 12 12 12 11 13 13 29 16 9 9 11 11 24 15 19 29 15 15 17 17
31290 13 24 14 12 11 12 12 12 11 13 13 29 16 9 9 11 11 24 15 19 29 15 15 17 17
33990 13 24 14 12 11 12 12 12 11 13 13 29
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 11 (Joseph Rice, 1800 - 1874 TN-AR)
Joseph 12 23 14 11 11 15 12 12 12 13 13 29 17 9 10 11 11 25 14 19 29 15 15 17 18
21928 12 23 14 11 11 15 12 12 12 13 13 29 17 9 10 11 11 25 14 19 29 15 15 17 18 >
30351 12 23 14 11 11 15 12 12 12 13 13 29 17 9 10 11 11 25 14 19 29 15 15 17 18
Group 12 (John G. W. Rice, c1807, Jefferson Co, TN)
John 13 22 14 10 13 15 11 14 11 12 11 28 16 8 9 8 11 23 16 20 28 12 14 15 16
21119 13 22 14 10 13 15 11 14 11 12 11 28 16 8 9 8 11 23 16 20 28 12 14 15 16
41527 13 22 14 10 13 15 11 14 11 12 11 28 16 8 9 8 11 23 16 20 28 12 14 15 16
Group 13 (Samuel R. Rice, 1744, Virginia)
Samuel 23 15 10 12 15 11 13 11 14 12 32 15 8 10 11 11 25 14 20 27 11 14 14 15
5106 15 23 15 10 12 15 11 13 11 14 12 31 15 8 10 11 11 25 14 20 27 11 14 14 15
6851 15 23 15 10 12 15 11 13 11 14 12 32
48918 16 23 15 10 12 15 11 13 11 14 12 32 15 8 10 11 11 25 14 20 27 11 14 14 15
49714 15 23 15 10 12 15 11 13 11 14 12 32 15 8 10 11 11 25 14 20 27 11 14 14 15
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 14 (Rices of Rices Landing, PA)
14295 13 22 14 10 14 14 11 16 11 12 11 28 15 8 9 8 11 23 16 20 28 12 14 15 16
47811 13 22 14 10 14 14 11 16 11 12 11 28 15 8 9 8 11 23 16 20 28 12 14 15 16 >
84143 13 22 14 10 14 14 11 16 11 12 11 28 15 8 9 8 11 23 16 20 28 12 14 15 16
84506 13 22 14 10 14 14 11 16 11 12 11 28 15 8 9 8 11 23 16 20 28 12 14 15 16
Group 15 (Rices of Rhode Island)
4507 13 25 14 11 11 13 12 12 12 13 14 29 17 9 10 11 11 25 14 18 30 15 16 16 17 >
86858 13 25 14 11 11 13 12 12 12 13 14 29 17 9 10 11 11 25 14 18 30 15 16 16 17 >
Group 16 (William Rice of Fauquier Co, VA)
William 13 24 14 10 12 15 13 12 12 13 13 29 18 9 10 11 11 25 15 19 29 15 15 16 18 >
4809 13 24 14 10 12 15 13 12 12 13 13 29 18 9 10 11 11 25 15 19 29 15 15 16 18 >
108045 13 24 14 10 12 15 13 12 12 13 13 29 18 9 10 11 11 25 15 19 29 15 15 16 18 >
Group 17 (Samuel Rice of Dedham, MA, and Woodstock, CT)
7648 13 22 14 10 13 14 11 14 11 12 11 28 15 8 9 8 11 24 16 20 28 12 14 15 16
111320 13 22 14 10 13 14 11 14 11 12 11 28 15 8 9 8 11 24 16 20 28 13 14 15 16
DYS
Locus:
3
9
3
3
9
0

1
9
3
9
1
3
8
5
a
3
8
5
b
4
2
6
3
8
8
4
3
9
3
8
9
i
3
9
2
3
8
9
ii
4
5
8
4
5
9
a
4
5
9
b
4
5
5
4
5
4
4
4
7
4
3
7
4
4
8
4
4
9
4
6
4
a
4
6
4
b
4
6
4
c
4
6
4
d
ID
Group 99 (other)
3145 13 24 14 10 11 15 12 12 14 13 13 29 17 9 10 11 11 26 15 19 30 15 15 17 17
21802